So, I went through again. I took one of the working 4MB modules from the motherboard. I verified it went missing. I then configured the jumpers on the memory module for 4MB. I tried each slot, actually, one by one. I fired up the amiga. I ran avail, whichamiga and showconfig each time. Each reported the CPU properly. All reported no change in RAM other than the missing 4MB dropping me down from 2MB CHIP + 16MB FAST to 2MB CHIP + 12MB FAST.
No dice. I double checked all the accelerator jumpers. None of those made a difference. I seated, reseated and repeated all the steps. It simply no longer seems to work.